We studied the deformation in a partially molten mantle through detailed petrostructural analysis of plagioclase lherzolites from the northwestern part of the central Lanzo massif. These peridotites display a pervasive anasto- mosed network of millimetre to centimetre-scale plagioclase-rich bands, which grades locally into a planar layering marked by an intercalation of plagioclase-rich and olivine-rich layers up to tens of centimetres wide. Both the anastomosed and the planar layering are subparallel to the deformation fabric, characterized by a foli- ation defined by the shape-preferred orientation of olivine and by crystal preferred orientation (CPO) of olivine, orthopyroxene, and clinopyroxene. This parallelism, the coexistence of microstructural evidence for melt–rock reactions and for deformation by dislocation creep, and the predominance of axial-[010] olivine CPO patterns, characterized by [010] axes clustered normal to the layering, indicate that deformation and the reactive melt per- colation that formed the layering were coeval. Strong heterogeneity in mineral composition in the planar layering domains, with Fe-enrichment in olivine and spinel and highly variable Ti content in spinel in cm-scale plagioclase-rich bands, implies that the latter interacted with higher melt volumes. High Ti contents in pyroxenes and spinel in the anastomosed layering domains point to changes in melt composition, hinting to less effective melt transport. We propose therefore that the planar layering records melt segregation in layers parallel to the shear plane, whereas the anastomosed layering results from melt alignment along grain boundaries subparallel to the shear plane, without segregation. Finite strain in the different layers cannot be quantified, but comparison of the present observations with data from experiments and from other mantle outcrops displaying shear- controlled melt organization suggests that the transition from the anastomosed to the planar layering might record an increase in finite strain, that is, strain localization associated with variations in the instantaneous melt fraction.