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Royal Society of Edinburgh, Transactions- Royal Society of Edinburgh, 03(94)

DOI: 10.1017/s0263593300000651

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The phylogenetic position of the extinct arachnid order Phalangiotarbida Haase, 1890, with reference to the fauna from the Writhlington Geological Nature Reserve (Somerset, UK)

Journal article published in 2003 by Jessica R. Pollitt, Simon J. Braddy, Jason A. Dunlop ORCID
This paper was not found in any repository; the policy of its publisher is unknown or unclear.
This paper was not found in any repository; the policy of its publisher is unknown or unclear.

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Abstract

ABSTRACTStudy of abundant phalangiotarbid (Arachnida: Phalangiotarbida) material – provisionally assigned here to Bornatarbus mayasii (Haupt in Nindel 1955) – from the Upper Carboniferous of Writhlington, UK, has revealed new information about some previously equivocal characters. The present authors report a trifurcate apotele, possible spiracles on sternite 5, and confirm the presence of 10 opisthosomal tergites plus a dorsal anal operculum. The affinities of phalangiotarbids are obscure, with most authors favouring affinities with Opiliones (harvestmen) and/or Acari (mites and ticks). Phalangiotarbida is scored for characters used in previous studies of arachnid relationships. A cladistic analysis based on 63 characters using 13 terminal arachnid taxa (plus a hypothetical outgroup), resolves Phalangiotarbida as sister group to (Palpigradi + Tetrapulmonata): the taxon Megoperculata sensu Shultz (1990). Even under cladistic analysis, the position of the Phalangiotarbida remains hard to resolve, but a prosomal sternite with distinct sclerites potentially groups them with the Megoperculata.