Published in
American Association for the Advancement of Science, Science, 6063(334), p. 1699-1703, 2011
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From Flat Foot to Fat Foot: Structure, Ontogeny, Function, and Evolution of Elephant "Sixth Toes"
Journal article published in 2011 by
John R. Hutchinson 
,
Cyrille Delmer,
Charlotte E. Miller,
Thomas Hildebrandt,
Andrew A. Pitsillides,
Alan Boyde
,
Cyrille Delmer,
Charlotte E. Miller,
Thomas Hildebrandt,
Andrew A. Pitsillides,
Alan Boyde
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Abstract
Elephants have an extra bone in their feet that can be traced through their fossil record to reveal a transition from flat to raised feet.