AbstractIn the frontal part of the Rhenohercynian fold-and-thrust belt (High-Ardenne slate belt, Germany), two successive types of quartz veins, oriented normal and parallel to bedding respectively, are interpreted to reflect the early Variscan compressional tectonic inversion of the Ardenne-Eifel sedimentary basin. Fracturing and sealing occurred in Lower Devonian siliciclastic multilayers under very low-grade metamorphic conditions in a brittle upper crust. A geometrical and microthermometric analysis of these veins has helped to constrain the kinematic and pressure-temperature conditions of both vein types, allowing the reconstruction of the stress-state evolution in a basin during tectonic inversion. It is demonstrated that bedding-normal extension veins, which developed under low differential stresses and repeatedly opened and sealed (crack-seal) under near-lithostatic fluid pressures, reflect the latest stage of an extensional stress regime. Bedding-parallel veins, which developed at differential stresses that were still low enough to allow the formation of extension veins, cross-cut the bedding-normal veins and preceded the regional fold and cleavage development. These veins show a pronounced bedding-parallel fabric, reflecting bedding-normal uplift and bedding-parallel shearing under lithostatic to supra-lithostatic fluid pressures during the early stages of a compressional stress regime. This kinematic history corroborates that fluid overpressures are easy to maintain during compressional tectonic inversion at the onset of orogeny.