Magnolia Press, Phytotaxa, 3(181), p. 138, 2014
DOI: 10.11646/phytotaxa.181.3.2
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Nine new species of Timonius (Rubiaceae) from Kinabalu Park are described: T. abanii, T. beamanii, T. bullatus, T. kinabaluensis, T. leopoldii, T. ophioliticus, T. pannosus, T. stenolobus, and T. tambuyukonensis. Of the nine species, six are likely to be endemic to the Kinabalu massif, and four appear to occur exclusively on ultramafic soils.